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74 TURTLES AND SNAKES




          A Specialized Mouth                                                                                                  Primitive Snakes


                                                                                                                                   Boas and pythons are called primitive, since they have neither
                                                                                                                                   fangs nor venom. These snakes have several rows of small,
                he most primitive snakes have heavy skulls and few teeth. Most snakes,                                         inward-curving teeth used for holding prey and swallowing quickly
                however, have lighter skulls and jointed jawbones. These joints are loose and                                  without letting the prey worm out. This feature is necessary for
          T can easily become dislocated so that the snake can swallow prey larger than                                        the snakes to be able to hold onto their prey, since they lack
                                                                                                                               venom. Venomous snakes, on the other hand, have little
          the natural shape of its own head. The teeth are fixed in the upper jaw or the                                       need to worry about their prey escaping, since they
          palate, and the fangs for injecting venom may be located either at the front                                         know that, after it is injected with the toxic
                                                                                                                               substance, it will not be able to get very far.
          or at the back of the mouth. Some species, as well as being large and
          powerful, have retractable fangs, allowing them to close their mouths
          when their fangs are not in use.                                                                                                                                                           Injecting Fangs
                                                                                                                                                                                                     Cobra species expel their venom in different ways
                                                                                                                                                                                                     depending on their fangs. The angle and direction of the
                                                                                                                                                                                                     opening determine how forcefully the liquid is injected.

          Cranial Anatomy
                                               JACOBSON'S ORGAN
              is directly related to each species' diet and—in  gives the snake an excellent sense
              the case of venomous snakes—to its system for  of smell. It consists of two
          injecting poison. Most snakes have small skulls with  cavities in the palate, into
          jawbones that can be separated voluntarily by sliding  which the snake brings its
          them along a kind of perpendicular rail, which  tongue after “tasting” the
          consists of a bone called the quadrate. This greatly  outside air. That is why
          increases the size of the snake's mouth.  snakes continually                                                                                                                               BLACK AND WHITE COBRA  BLACK-NECKED COBRA
                                               stick out their                                                                                                                                       Naja melanoleuca    Naja nigricolis
                                                                                                                                                                                                                         It has a jaw with a spur, but
                                               tongues.                                                                                                                                              It must bite its prey in order to
                                                                                                                                                                                                     inject venom.       it does not actually spit.
                                           DUVERNOY'S
                                           GLAND                                               RETRACTOR  FANGS
                                                                                               MUSCLE
                                                                                                            GLOTTIS
                                                                           UPPER JAW


           1    Viperid                                                                                                                                                                              ASIAN COBRA         RINKHALS SPITTING
                                                                                                                                                                                                     Naja naja
                                                                                                                                                                                                                         COBRA
                This type of skull has small                                                                                                                                                         It is the typical cobra, which  Hemachatus haemachatus
                teeth and large, retractable                                                                                    2    Colubrid                     3    Elapid                        bites to inject its venom.  It can spit great distances.
                fangs that are thick or                                                                                              This type of skull lacks front fangs.  In this type of skull, the fangs are
                hooked.                                                                                                              Some species are nonvenomous,     located in front, but they are smaller
                                                                                                                                     but others have fangs with a      and have only a groove, rather than
                                       QUADRATE                                                                                      groove for delivering venom.      a canal, for injecting venom.
                                                                                                                               Venom System                                                         Spitting Venom
                                                                                                                               consists of the two Duvernoy's glands, one on                        Forty cobra species can spray their venom
                                                                                                                               each side of the skull, which produce venom and  6.6 feet (2 m)      from a distance. They spray in self-defense
                                                                                                                               are connected to the fangs. When biting, muscle  THE DISTANCE FROM WHICH THE SPITTING   when they feel threatened. They can direct
                                                                                                                               contractions exert pressure on the gland and                         the stream into an enemy's eyes, causing
                                                                                                TEETH                          activate the injection mechanism.    COBRA CAN KILL BY SPRAYING ITS VENOM  grave damage or even death. The shape of
                                                           FUSED BONES                                                                                                                              their fangs is essential to this defense.

                                                                         The Deadliest Weapon                                                                                                           NON-SPITTING
                                                                         Rattlesnakes have long, thick fangs that are very sharp and                                                                    The long canal points
                                                                         kept folded inside their mouths. A movable joint at the base of                                                                downward and has a beveled
                                                                         the fang enables it to stand upright when the snake's mouth                                                                    edge at the end. The stream
                                                                         opens to bite.                                                                                                                 loses momentum.
                                                                         LENGTHWISE           CROSS SECTION
                                                                         The venom            The tooth has a                                                                                           SPITTING
                                                                         flows through  Entrance  cavity that                      SOLENOGLYPHS         PROTEROGLYPHS        OPISTHOGLYPHS              The canal's opening points
                                                                         the tube and         serves as the                    A   The hollow fangs are the  B  Small fangs in the front of  C  Fangs in the back, with no  forward and is narrow so
                                                                         directly into        canal for the                        only teeth in the jaw. They  the jaw, fixed in position  canal or groove. The prey  that it can eject the venom
                                                                         the prey.            poison.                              are long and retractable  and with a rear groove for  must be held in place.  more forcefully.
                                                                                                                                   and inject venom into the  conducting venom
                                                                                        Exit             Poison Canal              prey's tissues.
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