Page 79 - Reptiles & Dinosaurs (Britannica Illustrated Science Library)
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74 TURTLES AND SNAKES




 A Specialized Mouth  Primitive Snakes


           Boas and pythons are called primitive, since they have neither
           fangs nor venom. These snakes have several rows of small,
 he most primitive snakes have heavy skulls and few teeth. Most snakes,  inward-curving teeth used for holding prey and swallowing quickly
 however, have lighter skulls and jointed jawbones. These joints are loose and  without letting the prey worm out. This feature is necessary for
 T can easily become dislocated so that the snake can swallow prey larger than  the snakes to be able to hold onto their prey, since they lack
       venom. Venomous snakes, on the other hand, have little
 the natural shape of its own head. The teeth are fixed in the upper jaw or the  need to worry about their prey escaping, since they
 palate, and the fangs for injecting venom may be located either at the front  know that, after it is injected with the toxic
       substance, it will not be able to get very far.
 or at the back of the mouth. Some species, as well as being large and
 powerful, have retractable fangs, allowing them to close their mouths
 when their fangs are not in use.                                           Injecting Fangs
                                                                            Cobra species expel their venom in different ways
                                                                            depending on their fangs. The angle and direction of the
                                                                            opening determine how forcefully the liquid is injected.

 Cranial Anatomy
 JACOBSON'S ORGAN
 is directly related to each species' diet and—in  gives the snake an excellent sense
 the case of venomous snakes—to its system for  of smell. It consists of two
 injecting poison. Most snakes have small skulls with  cavities in the palate, into
 jawbones that can be separated voluntarily by sliding  which the snake brings its
 them along a kind of perpendicular rail, which  tongue after “tasting” the
 consists of a bone called the quadrate. This greatly  outside air. That is why
 increases the size of the snake's mouth.  snakes continually               BLACK AND WHITE COBRA  BLACK-NECKED COBRA
 stick out their                                                            Naja melanoleuca     Naja nigricolis
                                                                                                 It has a jaw with a spur, but
 tongues.                                                                   It must bite its prey in order to
                                                                            inject venom.        it does not actually spit.
 DUVERNOY'S
 GLAND  RETRACTOR  FANGS
 MUSCLE
 GLOTTIS
 UPPER JAW


 1  Viperid                                                                 ASIAN COBRA          RINKHALS SPITTING
                                                                            Naja naja
                                                                                                 COBRA
 This type of skull has small                                               It is the typical cobra, which  Hemachatus haemachatus
 teeth and large, retractable  2  Colubrid  3  Elapid                       bites to inject its venom.  It can spit great distances.
 fangs that are thick or  This type of skull lacks front fangs.  In this type of skull, the fangs are
 hooked.     Some species are nonvenomous,     located in front, but they are smaller
             but others have fangs with a      and have only a groove, rather than
 QUADRATE    groove for delivering venom.      a canal, for injecting venom.
       Venom System                                                         Spitting Venom
       consists of the two Duvernoy's glands, one on                        Forty cobra species can spray their venom
       each side of the skull, which produce venom and  6.6 feet (2 m)      from a distance. They spray in self-defense
       are connected to the fangs. When biting, muscle  THE DISTANCE FROM WHICH THE SPITTING   when they feel threatened. They can direct
       contractions exert pressure on the gland and                         the stream into an enemy's eyes, causing
 TEETH  activate the injection mechanism.   COBRA CAN KILL BY SPRAYING ITS VENOM  grave damage or even death. The shape of
 FUSED BONES                                                                their fangs is essential to this defense.

 The Deadliest Weapon                                                           NON-SPITTING
 Rattlesnakes have long, thick fangs that are very sharp and                    The long canal points
 kept folded inside their mouths. A movable joint at the base of                downward and has a beveled
 the fang enables it to stand upright when the snake's mouth                    edge at the end. The stream
 opens to bite.                                                                 loses momentum.
 LENGTHWISE  CROSS SECTION
 The venom  The tooth has a                                                     SPITTING
 flows through  Entrance  cavity that   SOLENOGLYPHS  PROTEROGLYPHS  OPISTHOGLYPHS  The canal's opening points
 the tube and  serves as the  A  The hollow fangs are the  B  Small fangs in the front of  C  Fangs in the back, with no  forward and is narrow so
 directly into  canal for the  only teeth in the jaw. They  the jaw, fixed in position  canal or groove. The prey  that it can eject the venom
 the prey.  poison.   are long and retractable  and with a rear groove for  must be held in place.  more forcefully.
          and inject venom into the  conducting venom
 Exit  Poison Canal  prey's tissues.
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