Page 12 - Fish and Amphibians (Britannica Illustrated Science Library)
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8 GENERAL CHARACTERISTICS                                                                                                                                                                               FISH AND AMPHIBIANS 9




          Earliest Forms                                                                                                                              Dunkleosteus                                 Dorsal


                                                                                                                                                      The Arthrodira—with a jointed   Its head was  fin                       The tail was not
                                                                                                                                                                                   protected by
                 bout 470 million years ago, the first fish appeared. Unlike today's                                                                  neck—were armored fish that  strong armor.                              protected by
                                                                                                                               STREAMLINED SHAPE      predominated in the late Devonian                                       scales.
                 fish, they did not have a jawbone, fins, or scales. Hard plates                                               The shape of Pteraspis  Period. The Devonian predator
          A covered the front part of the fish and formed a protective shield.                                                 shows that it was an   Dunkleosteus was an arthrodiran
                                                                                                                               excellent swimmer.     placoderm that lived over 300 million
          They also had a solid, flexible dorsal spine that allowed them to propel                                                                    years ago. Its head was encased in                                   This area of the body
          themselves. Later, in the Silurian Period, fish appeared that had a                                                                         an impressive set of plates 1.2 inches                               had neither armor
                                                                                                                                                      (3 cm) thick, with razor-sharp bony                                  nor scales.
          jawbone. Known as the gnathostomata, they were large predators.                                                                             plates that served as teeth.

                                                                                                                                                      FIERCE JAW                                                      It had a lobed tail, similar
                                                                                                                                                      Dunkleosteus was a                                              to a shark's tail, which
                                                                                                                                                      fierce predator that                                            indicates that it was a
                                                       CONICAL NOSE       EYES                                                                        devoured any type of                                            powerful swimmer.
          Pteraspis                                    Its streamlined shape  Very small,                                                             prey, including sharks.
                                                       helped the fish move.  located on both                                        DORSAL SPIKE
          The fish without a jawbone,                                     sides of the head.                                         Located on the fish's
          Pteraspis, was about 6.5                                                                                                   back, it worked like a
          inches (16 cm) long and lived in                                                                                           dorsal fin.                                                      It also had strong
          the seas of Europe, Asia, and North                                                                                                                                                         jaws with bony teeth.
          America. These fish were most
          abundant during the Devonian Period.
          They had bodies with armor that                                                                                                                DORSAL SPINES
          covered their heads, and they had a                                                                                                            These helped the fish
          streamlined shape. The shell had a co-                                                                                                         to stay balanced while
          nical nose that helped the fish to move.                                                                                                       swimming.
                                                                                                                                                                                                                     16 feet


                6.5 inches (16 cm)                                                                                                                                                                                   (5 m)
          WING SHIELD
                                                                                                                                                                                                                      LENGTH OF THE FISH
          Scientific Pteraspis
          name
                                      MOUTH
          Diet   Small organisms      Having no jawbone,                                                                                                                                                              TAIL
                                                                                                                                                                                                                      The shape of the tail
          Habitat   Sea, then rivers and lakes  they fed on small                                                                                                                                                     helped balance the
                                      organisms.
          Range  Europe, Asia, North America                                                                                                                                                                          weight of the armor.
          Period  Early Devonian
          EVOLUTION OF THE JAWBONE cranium
                                                                                                                                                      LATERAL LINE
          The development of the jawbone was a  The evolution of the jawbone
          long evolutionary process that involved  modified the configuration                                                                         Sensory organs are
          changes in the diet of fish to include not  of the skull.                                                                                   present on both sides
          only small organisms but also other fish.                                                                                                   of the body and on top
                                                                                                                                                      of the armor.
             PRIMITIVE
           1
             VERTEBRATE
                                                                     Marine
             The first fish
                                                                     lamprey                                                                                               Chimaeriformes              Cheirolepis   Pycnodus      Sole
             had no jawbone.
                                                                                  Fossil
                                                                                      Fish with lungs appeared
                                                                                      in the Mesozoic Era                           Evolution                               Holocephali              Cheirolepididae  Pycnodontiformes  Holostei Teleostei
             ELASMOBRANCHIMORPH                                                   (200 million years ago).                                                  Wing shield
           2                                                                                                                                                         Dunkleosteus  Sharks and rays  Eusthenopteron  Chondrostei  NEOPTERYGII
             The formation of the                                                 Similar to amphibians, these                          In the Devonian Period  (Pteraspis)
             jawbone permitted new                                   Placoderms   species breathe with lungs                            ocean fish began to
             feeding habits, and the                                              and are now considered                            diversify. Coelacanths appeared,  Placoderms  Elasmobranchii  Sarcopterygii
             fish evolved from                                                    living fossils. The line                          as well as the earliest bony fish  Lamprey
             herbivore to carnivore.                                              through the center of the                         and the first cartilaginous fish,                                ACTINOPTERYGII
                                                                                                                                                                                       Acanthodii
                                                                                  photo of the fossil is the                        including sharks. In this period
                                                                                  fish's lateral line.                              the three main groups of  Jawless      CARTILAGINOUS FISH   BONY FISH
                                                                                                                                                              fish
                            BONY FISH                                                                                               gnathostomad fish also                                                    Devonian
                         3
                            They already had a                                                                                      appeared: the placoderms,
                                                                                  FOSSILIZED                                                                        GNATHOSTOMATA
                            specialized jawbone                      Modern fish  LUNGFISH SCALES                                   chondrichthyes, and                                                        THIS PERIOD SAW AN EXPLOSION IN
                            like fish of today.                                   Dipterus valenciennesi                            osteichthyes.          VERTEBRATA                                          THE DIVERSITY OF FISH SPECIES.
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