Page 8 - (DK) Ocean - The Definitive Visual Guide
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               About this Book                                               Ocean Environments

                                                                            This chapter looks at specific parts of the oceans. It is divided into sections
               THIS BOOK IS DIVIDED INTO four chapters. An overview         on different zones, starting with coasts and the seashore and then
                                                                            moving to progressively deeper waters, first with shallow seas and then
               of the physical and chemical features of the oceans is
                                                                            the open ocean and ocean floor. A final section, polar oceans, looks
               given in the introduction; ocean environments                at the frozen waters around the North and South Poles. In each section,
               looks at the main zones of the oceans, and ocean life        explanatory pages describe typical features and formative processes, while
                                                                            the succeeding pages contain profiles of actual features. The profiles are
               examines the life-forms that inhabit them. The atlas         arranged by geographical location, starting with
               of the oceans contains detailed maps of the oceans.          the Arctic Ocean and followed, in order, by the   160  SHALLOW SEAS
               Most chapters are divided into smaller sections.             Atlantic, Pacific, Indian, and Southern Oceans.  Shiraho Reef  species concentrated in a few square   Tubbataha Reefs
                                                                                                                                           PACIFIC OCEAN WEST
                                                                                                                            PACIFIC OCEAN WEST
                                                                                                                                   kilometres. The reef also contains the
                                                                                                                                   world’s largest colony of rare Blue
                                                                                                                               TYPE  Fringing reef  Ridge Coral (Heliopora coerulea). For   TYPE  Atol
                                                                                                                                   decades, environmentalists battled
                                                                                                                               AREA  10 square km    to save the reef from the building of    AREA  330
                                                                                                                               (4 square miles)  a new airport for Ishigaki. A proposal   (130 squar
                                                                                                                               CONDITION Reasonable;   to construct the airport on top of the   CONDITION
                                                                                                                               damaged in parts by   reef was dropped, but concern remains   recovering
                                                                                                                               bleaching in 1998, 2007  now that it has been built on land, as   bleaching
                                                                                                                            153
                                                                            152  SHALLOW SEAS                           CORAL REEFS LOCATION  Southeast coast of Ishigaki Island, at the   discharge of excavated soil into the   LOCATION  Central Sulu Sea, between t
                                                                                                                           southwestern extremity of Japanese archipelago  and northern Borneo
                                                                                            CORAL DIVERSITY                        reef is likely to have an adverse effect.
                                                                              Coral Reefs   In this seascape off a Fijian island,    Reef Formation  BLUE RIDGE CORAL  The Tubbataha Reefs lie arou
                                                                                            groups of shoaling sea goldies hover
                                                                                                              The individual animals that make up corals are called polyps. The polyps of the main group  Shiraho Reef, off Ishigaki Island, part
                                                                                            over diverse species of coral, sponges,
                Introduction                                                  CORAL REEFS ARE SOLID STRUCTURES built from the remains    The polyps also form colonies that create community skeletons in a variety of shapes. An   notice in the 1980s as an outstanding   varies from violet through blue, turquoise,    are famous for the many larg
                                                                                                                                   Despite its name, the colour of this coral
                                                                                                                                           atolls in the centre of the Sul
                                                                                            and other reef organisms.
                                                                                                              of reef-building corals, stony corals, secrete limestone, building on the substrate underneath.  of the Japanese archipelago, came to
                                                                                                                                           (open ocean) marine animals
                                                                              of small marine organisms, principally a group of colony-forming
                                                                                                              important contributor to the life of these corals is the presence within the polyps of tiny
                                                                                                                           example of biodiversity, with some
                                                                                                                                   and green to yellow-brown. Its branching
                                                                                                              organisms called zooxanthellae, which provide much of the polyps’ nutritional needs.
                                                                              animals called stony (or hard) corals. Reefs cover about 108,000
                                                                                                                                           turtles, and barracuda. The ste
                                                                                                                    Grazing and boring organisms also contribute, by
                                                                              square miles (280,000 square km) of the world’s shallow marine   Other organisms that add their skeletal remains to the reef include mollusks and echinoderms.  120 species of coral and 300 fish   vertical plates can form massive colonies.  to them – such as sharks, Ma
                                                                              areas, growing gradually as the organisms that form their living   breaking coral skeletons into sand, which fills gaps   shelving reefs here are also ri
                                                                                                                    in the developing reef. Algae and other encrusting
                                                                              surfaces multiply, spread, and die, adding their limestone skeletons   organisms help bind the sand and coral fragments   smaller life, including many s
               This opening chapter is divided into four sections. In ocean water,   to the reef. Coral reefs are among the most complex and beautiful   together. Most reefs do not grow continuously but   of crustaceans, colourful nud
                                                                                                                    experience spurts of growth interspersed with
                                                                              of Earth’s ecosystems, and are home to a fantastic variety of animals
                                                                                                                                           (sea slugs), and more than 35
                                                                              and other organisms; but they are also among the most heavily
                                                                                                                    quieter periods, which are sometimes associated
                                                                                                                                           of stony and soft coral.
                                                                              utilized and economically valuable. Today, the world’s reefs are   with recovery from storm damage.  STONY CORAL  Reefs were rated by scuba di
                                                                                                                                           In the early 1990s, the Tub
                                                                              under pressure from numerous threats to their health.
               the properties of water itself are examined. ocean geology covers the                                    branching hard     among the top ten dive sites
                                                                                                                        This group of
                                                                                                                        corals is growing
                                                                              Types of Reefs
                                                                                                                                           world. However, during the
                                                                                                                        at a depth of about
                                                                                                              At the center of each polyp is
                                                                                                                        coast of eastern
                                                                              Coral reefs fall into three main types: fringing reefs, barrier reefs, and    OPEN POLYPS  16 ft (5 m) off the   they suffered considerable da
                                                                                                                                           from destructive fishing pract
                                                                                                                        Indonesia. Individual
                                                                                                              an opening, the mouth, which
               materials of the ocean floor and the way that it changes over time.   with little or no separation from the shore, and develop through upward   around the gut secretes limestone,   grow up to a few   the establishment of a seawee
                                                                              atolls. The most common are fringing reefs. These occur adjacent to land,
                                                                                                              leads to an internal gut. The tissue
                                                                                                                        stony corals can
                                                                              growth of reef-forming corals on an area of continental shelf. Barrier reefs
                                                                                                                        inches per year.
                                                                                                              which builds the reef.
                                                                              are broader and separated from land by a stretch of water, called a lagoon, that
                                                                              can be many miles wide and dozens of yards deep.             CORAL DROP-OFF
                                                                              Atolls are large, ring-shaped reefs, enclosing a central lagoon; most atolls    Distribution of Reefs  In this photograph of a steeply she
                                                                              are found well away from large landmasses, such as in the South Pacific. Parts
               circulation and climate is about the interaction between oceans    of the reef structure in both atolls and barrier reefs often protrude above sea   at least 64˚F (18˚C), and preferably 77–84˚F (25–29˚C). They grow best where the   reef slope, several species of soft
                                                                                                               Stony corals can grow only in clear, sunlit, shallow water where the temperature is
                                                                                                                                           are visible, together with a shoal o
                                                                                                               average salinity of the water is 36 ppt (parts per thousand) and there is little wave
                                                                                  level as low-lying coral islands—these develop as
                                                                                                                                           Longfin Bannerfish.
                                                                                                               action or sedimentation from river runoff. These conditions occur only in some
                                                                                  wave action deposits coral fragments broken off
                                                                                  from the reef itself.        tropical and subtropical areas. The highest concentration
                                                                                  Two other types of reefs are patch reefs—small   of coral reefs is found in the Indo-Pacific region,
                                                                                                               which stretches from the Red Sea to the central
                                                                                  structures found within the lagoons of other reef
               and the atmosphere and the large-scale movement of water, while    types—and bank reefs, comprising various reef   Pacific. A smaller concentration of reefs occurs   PACIFIC OCEAN WEST  Nusa Tenggara is a chain of around    indicate an extremely high d
                                                                                                                                           marine life in this region. Fo
                                                                                  structures that have no obvious link to a coastline.
                                                                                                                                   500 coral-fringed islands in southern
                                                                                                               around the Caribbean Sea. In addition to
                                                                                                               warm-water reefs, awareness is growing about
                                                                                                               other corals that do not depend on sunlight,   Nusa Tenggara  Indonesia. The northern islands are   a single large reef can contai
                                                                                                               and form deep, cold-water reefs—some of   TYPE  Fringing reefs,   volcanic in origin, while the southern   than 1,200 species of fish (m
               tides and waves looks at movements and disturbances of water                                        WARM-WATER    barrier reefs  islands consist mainly of uplifted coral   in all the seas in Europe com
                                                                                                               them outside the tropics (see p.178).
                                                                                                                                   limestone. Many of the reefs have
                                                                                                                                           and 500 different species of
                                                                                                                   REEF AREAS
                                                                                                                               AREA  5,000 square km
                                                                                                                   The conditions needed for   (2,000 square miles)  been only rarely explored. However,   building coral. Common ani
                                                                                                                   the growth of warm-water   CONDITION  Damaged by   what surveys have been carried out   include Eagle Rays, Manta R
               on a smaller scale.                                            FRINGING REEF  BARRIER REEF  ATOLL An atoll is a ring of coral reefs or coral   within tropical areas of the   COLD-WATER CORAL  LOCATION  Southern Indonesia, from Lombok in the
                                                                                                                   coral reefs are found mainly
                                                                                                                               fishing practices
                                                                                                                   Indian, Pacific, and Atlantic
                                                                                  A barrier reef is separated from the coast by a
                                                                              A fringing reef directly borders the
                                                                                       islands enclosing a central lagoon. It
                                                                              shore of an island or large
                                                                                  lagoon. In this aerial view, the light blue area is the
                                                                                                                   oceans. The reefs are chiefly
                                                                                                                   in the western parts of these
                                                                              landmass, with no deep lagoon.
                                                                                  reef and the distant dark blue area is the lagoon.
                                                                                       may be elliptical or irregular in shape.
                                                                                                                   oceans, where the waters   This species, Lophelia pertusa, is one of a few  west to Timor in the east
                                                                                     island subsides               are warmer than in the   of the reef-forming corals that grow in cold
                                                                              coral grows on  sea level  when volcano has
                                                                              shoreline, forming  become inactive  eastern areas.  water, at depths to 1,650 ft (500 m).
                                                                              fringing reef   BARRIER REEF
                                                                                   FRINGING REEF                 HUMAN IMPACT
                                                                                                                 CORAL BLEACHING
                                                                                                                 reef-building corals and occurs
                                                                                                                 when the tiny organisms called
       32  OCEAN WATER                    THE CHEMISTRY OF SEAWATER  33  ◀ OCEAN WATER  OCEAN ENVIRONMENTS  ATOLL FORMATION  lagoon of  lagoon  reef face  coral continues   Bleaching refers to color loss in
                                                                                                                 zooxanthellae, which give corals
                                                                              volcanic
                                                                                                                 their colors, are ejected from coral
                                                                              island
                                                                                   shallow water
                                                                                          to grow, forming
                                                                                                                 polyps or lose their pigment. In
                                                                                          barrier reef
        The Chemistry of Seawater  volcanic ash drifts   SOURCES, SINKS, AND EXCHANGES  KEY  gases  Gases in Seawater  An atoll is shown here forming around a   ATOLL  volcanic island  extreme cases, this can lead to the   OCEAN ENVIRONMENTS
                               Shown here are various sources, sinks,
                               and exchange processes for the ions,
                                                                                                                 coral’s death. Various stresses
                                                                              volcanic island. First, the island’s shore is
                                      The main gases dissolved in seawater are nitrogen (N), oxygen (O 2 ),
                         down to sea  salts, and minerals (yellow arrows),   ions, salts, and   and carbon dioxide (CO 2 ). The levels of O 2  and CO 2  vary in response    colonized by corals forming a fringing reef   becomes submerged  can cause bleaching, including
                                    minerals
                               (turquoise arrows) in seawater.
        THE OCEANS CONTAIN MILLIONS OF DISSOLVED chemical substances. Most of   spread of  volcanic ash   gases (pink arrows), and plant nutrients   plant nutrients  to the activities of photosynthesizing organisms (phytoplankton) and   This section covers the properties   (above). Over time, the island subsides, but   central area filled  pollution, ocean temperature rise,
                                                                                                                 and ocean acidification (see p.67).
                              and gases into
                                                                              coral growth continues, forming a barrier
        these are present in exceedingly small concentrations. Those present in significant   rain clouds  animals. The level of O 2  is generally highest near the surface, where the   reef (above right). Finally, the island   by reef limestone  In recent decades, some mass
                                      gas is absorbed from the air and also produced by photosynthesizers.
        concentrations include sea salt, which is not a single substance but a mixture   Its concentration drops to a minimum in a zone between about 660 ft    disappears, but the coral maintains growth,   coral continues   bleaching events have affected
        of charged particles called ions. Other constituents include gases such   (200 m) and 3,300 ft (1,000 m), where oxygen is consumed by bacterial   of the water molecule, the   forming an atoll (right). Atolls can also form   to grow where   reefs over wide areas.
                                                                              as a result of sea-level rise.
                                                                                         waves bring food
                                      oxidation of dead organic matter and by animals feeding on this matter.
        as oxygen and carbon dioxide. One reason the oceans contain so   Deeper down, the O 2  level increases again. CO 2  levels are highest at depth
        many dissolved substances is that water is an excellent solvent.
                                      and lowest at the surface,
                                      where the gas is taken up by
        The Salty Sea  salts are leached   photosynthesizers faster than   chemistry of seawater, and
                  from rocks into rivers
                                      it is produced by respiration.
        The salt in the oceans exists in the form of charged   and streams and    salt spray
                  flow to ocean
                      onto land
        particles, called ions, some positively charged and some   CARBON SINK
        negatively charged. The most common of these are sodium   nutrients from    Many marine animals, such as
                 soil wash into
                                      nautiluses (below), use carbonate
        and chloride ions, the components of ordinary table salt   rivers and streams,    (a compound of carbon and oxygen)
                 and flow to ocean
        (sodium chloride). Together they make up about 85 percent   in seawater to make their shells.
        by mass of all the salt in the sea. Nearly all the rest is made   washing of ions   After they die, the shells may form   the way that attributes such    OCEAN ENVIRONMENTS
                                    from volcanic dust
        up of the next four most common ions, which are sulfate,   and gases into sea,   sediments and eventually rocks.  ▲ EXPLANATORY PAGES
                                    dissolved in rain
        magnesium, calcium, and potassium. All these ions, together
        with several others present in smaller quantities, exist   dust blown off land
        throughout the oceans in fixed proportions. Each is
        distributed extremely uniformly—this is in contrast to   exchange of gases   as temperature, pressure,
        some other dissolved substances in seawater, which are   exchange of gases   between ocean and   These pages describe general types of
        unevenly distributed.    between animals    atmosphere                                                             REEF FLAT OFF PANTAR ISLAND
                                 and seawater
                                           OXYGEN PRODUCER AND CONSUMER                                                    This shallow reef area, featuring
           BREAKDOWN OF SALT               Oxygen levels in the upper ocean depend
           If 2½ gallons (10 liters) of seawater are   on the balance between its production by                            numerous species of stony coral and
           evaporated, about 12 3 / 4 oz (354 g) of salts   photo-synthesizing organisms, such as kelp,    and light transmission change    a starfish, is in east Nusa Tenggara.
                                           and its consumption by animals, such as fish.
           are obtained, of the types shown below.                          environments. The example above is
           2½ gallons (10 liters) of seawater
                                      Nutrients
            other salts  1 /4oz (7.5g)  Numerous substances present in small amounts in seawater are essential   with depth in ocean water.
             calcium sulfate                                                taken from the SHALLOW SEAS section.
             (gypsum)  2 /3oz (17.7g)  for marine organisms to grow. At the base of the oceanic food chain are
             magnesium salts   +
             2oz (54.8g)  +  +  sodium   ion (positive   phytoplankton—microscopic floating life-forms that obtain energy by
                                      photosynthesis. Phytoplankton need substances such as nitrates, iron, and
             sodium chloride   –  charge)
             (halite)    –  –         phosphates in order to grow and multiply.  If the supply of these nutrients
             10oz (274g)  +  –  Na +  –  +  slow uplift of   dries up, their growth stops; conversely, blooms (rapid growth phases)
                  sedimentary rocks
                – –  –  at continental   occur if it increases. Although the sea receives some input of nutrients
               +  +  margins, exposing   from sources such as rivers, the main supply comes from a continuous
         WATER AS A SOLVENT  +  salts, minerals,   uptake of   nutrient   cycle within the ocean. As organisms die, they sink to the ocean floor,
                  and ions at surface
                    nutrients by
         The charge imbalance on its   phytoplankton  upwelling  where their tissues decompose and release nutrients. Upwelling of seawater
         molecules makes water a good   –
         solvent. When dissolving and   –  –  exchange of    from the ocean floor (see p.60)
                       gases between
         holding sodium chloride in   +  phytoplankton    recharges the surface waters with
         solution, the positive ends of   +  +  Sources and Sinks  and seawater  sinking and   vital substances, where they are
         the molecules face the chloride   sodium chloride   –  +  Cl –  +  –  The ions that make up the salt in the oceans have arrived   decomposition   release of minerals   taken up by the phytoplankton,
                                   from hydrothermal
             crystal
         ions and the negative ends face   + +  +  there through various processes. Some were dissolved out of   of dead   vents  refueling the chain.
         the sodium ions.  water molecule  –  –  rocks on land by the action of rainwater and carried to the   organisms
             chloride ion        dissolving of   PLANKTON BLOOM
             (negative charge)  –  sea in rivers. Others entered the sea in the emanations of hydrothermal   minerals from   This satellite image of the
                 vents (see p.188), in dust blown off the land, or came from volcanic ash.   sea floor  Skagerrak (a strait linking the North
        PEOPLE   There are also “sinks” for every type of ion—processes that remove them   precipitation of minerals    and Baltic seas) shows a bloom of
                                      phytoplankton, visible as a turquoise
        ALEXANDER MARCET  from seawater. These range from salt spray onto land to the precipitation   onto sea floor   discoloration in the water.
                 of various ions onto the seafloor as mineral deposits. Each type of ion
        The Swiss chemist and doctor Alexander   has a characteristic residence time.   carbonates   SILICEOUS DIATOMS
                              incorporated
        Marcet (1770–1822) carried out some of the   This is the time that an ion remains   into seafloor   These tiny forms of    Ocean Life
                               planktonic organisms
                              sediments from
        earliest research in marine chemistry. He is
       INTRODUCTION  the main chemical ions in seawater (such as   have long residence times, ranging   42  OCEAN GEOLOGY  INTRODUCTION  THE ORIGIN OF OCEANS AND CONTINENTS  43
                 in seawater before it is removed.
                               have cell walls made
                              animal shells
        best known for his discovery, in 1819, that all
                 The common ions in seawater
                               of silicate. They can
        sodium, chloride, and magnesium ions) are
                               only grow if there are
                 from a few hundred years to
        present in exactly the same proportions
                               sufficient amounts
                               of silica present
        throughout the world’s oceans. The
                 hundreds of millions of years.
                               in the water.
        unchanging ratio between the ions holds
        true regardless of any variations in the
                 RIVER DISCHARGE
                 River discharge is a mechanism by which ions
        salinity of water and is known
                 of sea salt and nutrients enter the oceans.
        today as the principle of
                 Here, the Noosa River empties into the sea on
        constant proportions.
                 the coast of Queensland, Australia.
                               The Origin of Oceans    DEVELOPMENT OF CONTINENTAL CRUST  Water and Atmosphere  BANDED IRON  This chapter contains two sections. The introduction to ocean life
                                                    Modification of the crust above rising mantle
                                                                   Known as a banded-iron
                                                    flows was delayed by the continuous intrusion
                                                                   formation, this layered rock
                                                        During the process of differentiation, volatile
                                                    of mantle basalt, resulting in the greenstone
                                                                   formed as the oxygen content
                                                    belts found today at the heart of each   materials were expelled from Earth’s interior    contains iron oxides that
                               and Continents  zircon crystals,   greenstone belts above   hydrogen and helium, would quickly have been    rivers erode and   covers the ecology and history of marine life and the way that marine
                                                                   of early oceans increased.
                                                        by volcanic activity. The lightest gases, such as
                                                    continental shield.
                                                        lost to space, leaving a stable atmosphere of
                                                    rising mantle flow
                                                                    transport sediment
                                                        nitrogen, carbon dioxide, and water vapor. Some
              OCEAN GEOLOGY ▶  EARTH’S OCEANS FORMED MORE THAN 4 billion years ago, mainly from    continental crust  ZIRCON  primitive   basalt  continuously   crust pulled apart   of the water vapor would have condensed to form
                                            among the earliest
                                            materials
                               water vapor that condensed from its primitive atmosphere but also from water
                                                        liquid water, and it seems there was a significant
                                                    intrudes from
                                                        ocean earlier than 4 billion years ago.
                                                      by convective
                                                    mantle
                                                      motion in mantle
                               brought from space by comets. Initially, after acquiring a layered internal structure,
                                                        Some meteorites contain 15-20 percent
                                               thickens above
                               the Earth had a uniform crust that was enriched in lighter elements and floated   continental crust   water and the early Earth is thought    ocean water from
                                                              volcanic eruptions
                               on an upper mantle made of denser materials. Later, the crust became   sinking mantle   to have had the same composition,    and comet   organisms are classified. It is followed by a larger section, kingdoms of
                                                              impacts
                                               flow, without
                                                         providing an ample source for the
              As well as describing   differentiated into two types as continents began to form, made from    mantle  interference  early ocean. More water arrived
                               rocks that were chemically distinct from those underlying the oceans.
                                                         with impacting comets. It was in
                                                         the ocean that free oxygen first
                               Continental Crust  sedimentary   appeared, with the arrival    traces of early
              the composition of the   The continents include a wide range of rock    THE OLDEST ROCKS  rocks  around 3.5 billion years ago.   meteorite    ocean life. This is divided into domains or kingdoms and, in the case
                                                          of photo- synthesizing life
                                                              and comet
                                     These sedimentary
                                                              bombardment
                               types, including granitic igneous rocks, sedimentary
                               rocks, and the metamorphic rocks formed by the   rocks on Baffin Island   primitive   gradually erased
                                            oceanic
                               alteration of both. They contain a lot of quartz, a   lie on the Canadian   crust
                                     Shield. The stable
                               mineral absent in oceanic crust. The first continental   continental shields   THE EARLY EARTH
             ocean floor, this section     rocks were the result of repeated melting, cooling,   contain the world’s  mantle  Earth had deep oceans from    of the plant and animal kingdoms, further divided into smaller groups.
                                                             an early stage, with volcanoes
                               and remixing of oceanic crust, driven by volcanic
                                     most ancient rocks,
                                     which are around
                               activity above mantle convection cells, which were
                               much more numerous and vigorous than today’s.   4 billion years old.  vigorous convection   and an increasing area of
                                                             continental crust standing
                                                          cells in upper mantle
                               Each cycle left more of the heavier components in   above the surface. The ocean   rifts occur when
                                                             became salty as weathering
              looks at the processes   the upper mantle and concentrated more of the lighter components in   of surface rocks added    fragments of crust   volcanic
                                                                   eruptions
                                                                move apart
                               the crust. The first microcontinents grew as lighter fragments of crust
                                                             minerals to the water.
                                                                   add gases and
                                                                   water vapor to
                               collided and fused. Thickening of the crust led to melting at its base
                               and underplating with granitic igneous rocks. Weathering accelerated   atmosphere  In each case, a general overview of the organisms that make up the
                               the process of continental rock formation, retaining the most resistant   volcanic activity   liquid outer core
                                         adds igneous rocks
                               components, such as quartz, while washing solubles into the ocean.  to surface above
               that shape it, tracing    basaltic  lava  rift  basalt sheets
                                         rising flows
                                 (dikes)
                                  sediment
                                   ocean   Oceanic Crust
                                   surface  The oceanic crust has a higher density than the   solid inner core  group is followed by profiles of a selection of individual species. The
             the origin of the oceans   ocean   continental crust, making it less buoyant. Both types
                                    of crust can be thought of as floating on the “plastic”
                                    upper mantle, and the oceanic crust lies lower due to
                              gabbro  crust  its lower buoyancy. It is relatively thin, with a depth
                               peridotite  of never more than 7 miles (11 km), compared with a
                                    thickness of 15–43 miles (25–70 km) for most continental   section begins with the smallest forms of life, the bacteria and archaea,
                and their changing     lithosphere crust. It consists mainly of basalt, an igneous rock that is
                                    low in silica compared with continental rocks, and richer
                               Moho  in calcium than the mantle. Basalt lava is created when
                                    hot material in the upper mantle is decompressed, allowing
                                 magma  top layer   it to melt and form liquid magma. The decompression
                               asthenosphere  rises to  surface  of upper  mantle   occurs beneath rifts in the crust, such as those found at the
               size and shape over   OCEAN-FLOOR STRUCTURE  mid-ocean ridges, and it is through these rifts that lava is   and ends with the animal kingdom.
                                    extruded onto the surface to create new ocean crust.
                               Three layers of basalt in the crust (basaltic
                               lava, dikes, and gabbro) are separated from
                               the mantle by the Mohorovic ˇic ´ discontinuity
                               (the Moho). The top layer of the upper mantle
                 geological time.  rigid lithosphere, which makes up tectonic
                               is fused to the base of the crust to form the
                               plates.The asthenosphere is the soft zone
                               over which the plates of the lithosphere glide.
                              INTRODUCTION  MANTLE ROCKS          ANDEAN VOLCANOES  INTRODUCTION
                                Peridotite is the dominant rock type
                                found in the mantle, consisting of
                                silicates of magnesium, iron, and
                                                                  This radar image shows volcanoes
                                other metals. Sometimes it is
                                                                  formed from andesite lava, whose
                                brought to the surface when parts
                                                                  oceanic and continental rocks.
                                here in Newfoundland, Canada, or
                                as fragments from volcanic activity.
       54  CIRCULATION AND CLIMATE  of the ocean floor are uplifted, as   55  composition is intermediate between   DOMAIN  Eucarya
        Ocean Winds  polar-front jet stream—narrow  polar easterly  polar cell  air rises in   LONG-HAUL SAILING
                       subpolar
                       latitudes
                              Winds can blow with a consistent
                              strength and direction over large areas of
                 ribbon of strong wind at high   Ferrel cell
        THE PATTERN OF AIR MOVEMENT over the oceans   altitude at top of front  air descends   ocean. Consequently, on long-haul sailing
                              trips, the same basic sail settings can
        results from solar heating of the atmosphere and    southwesterly   in subtropical   often be used for days on end.
                        latitudes
                  wind
        Earth’s rotation. This pattern of winds is modified by   Hadley
        linked areas of low and high pressure (cyclones and   cell                 KINGDOM  Animalia
        anticyclones), which continually move over the oceans’   air rises                               276  ANIMAL LIFE
        surface. Near coasts, additional onshore and offshore   at equator  ◀ CIRCULATION AND CLIMATE
        breezes are common. These are caused by differences
        in the capacity of sea and land to absorb heat.
               direction of                                                                                  Mollusks
        Atmospheric Cells  Earth’s spin            This section describes the       PHYLUM  Mollusca
        Solar heating causes the air in Earth’s atmosphere to   CIRCULATION CELLS
        cycle around the globe in three sets of giant loops,   The atmospheric cells
               produce north–south
        called atmospheric cells. Hadley cells are produced by   airflows. These are   north-
        warm air rising near the equator, cooling in the upper   modified by Earth’s   easterly                DOMAIN  Eucarya   AMONG THE MOST SUCCESSFUL of all marine animals,
               spin, producing winds
        atmosphere, and descending to the surface around   that blow diagonally.  trade  wind  large-scale circulation of the
        subtropical latitudes (30˚N and S). Then the air moves                                                     mollusks display great diversity and a remarkable range
        back toward the equator. Ferrel cells are produced by   subtropical   southeasterly                  KINGDOM  Animalia
                        trade wind
                 jet stream
        air rising around subpolar latitudes (60˚N and S), cooling and falling in   polar-front jet   CLASSES  8   of body forms, allowing them to live almost everywhere
        the subtropics, and then moving toward the poles. Polar cells are caused   stream                     PHYLUM  Mollusca
        by air descending at the poles and moving toward the equator.  air descends    trade winds meet at Intertropical
                   at pole  Convergence Zone       oceans, both deep down and                                      from the ocean depths to the splash zone. They include
        initial direction   DISCOVERY                                                                         CLASSES  8
        of air   The Coriolis Effect
        movement  SATELLITE IMAGING                                                                                oysters, sea slugs, and octopuses. Most species have shells
           air   The atmospheric cells cause north–south                                                      SPECIES  73,683
           deflected
           to right  air movements. These are altered by the  Ocean winds are monitored                            and are passive or slow-moving; some lack eyes. Others
            Coriolis effect. As the Earth spins, parcels of   by instruments called  at the surface. It also looks at   SPECIES  73,683
            air at different latitudes in the atmosphere   scatterometers, such
           have different west-to-east velocities (air at   as an instrument                                 are intelligent, active hunters with complex nervous systems and large eyes.
           the Equator moves fastest). When they change   called ASCAT on  air ascends    Pressure-system Winds
                               from cyclone
            latitude by moving to the north or south, they   the METOP-A  warm air rising                    Filter-feeding mollusks, such as clams, are crucial to coastal ecosystems,
           retain these west-to-east velocities, which differ   satellite (right).  air descends into   In any area of ocean where air sinks—often at subtropical latitudes—a zone of   ocean climates and the many
                 A scatterometer
                                     high atmospheric pressure, or anticyclone, develops. Where warm air rises, areas
              from those of air in the
                                   anticyclone
           AIR DEFLECTIONS  is a radar device  low pressure                                                  as they provide food for other animals and improve water quality and clarity.
           In the Northern Hemisphere,   latitudes they move into.   that can measure  ASCAT antenna   at center  central area of   of low pressure, called cyclones or depressions, occur. These often develop near
                                   high pressure
              Hence, the air veers to
                                     the equator and subpolar latitudes. Cyclones and anticyclones create linked,
           the Coriolis effect causes   both wind speed  (one of three)
           all air movements to be   the east (in the direction   and direction.  cold air sinks   circulating wind patterns, which continually move and change. In the Northern   Many mollusks are commercially important for food, pearls, and their shells.
         Earth’s   air   deflected to the right of   of Earth’s spin) when   Hemisphere, there is a clockwise movement of air
         rotation  deflected   their initial direction. In    moving away from the   around an anticyclone, and a counterclockwise motion   ways that the oceans and the
        initial direction    to left  the Southern Hemisphere,   Equator and to the west  Prevailing Winds  CYCLONES AND   around a cyclone. This pattern is reversed in the
                                     ANTICYCLONES
                                     Air moves from an area of
           they veer to the left.
        of air movement  when moving toward it.  The winds produced by pressure differences and modified by the   air spirals  around central   high pressure toward one    Southern Hemisphere. Local pressure systems can affect
                                        the general pattern of prevailing winds. In particular,
               northeasterly   area of low   of low pressure, but the                                        Anatomy
            polar   monsoon   Coriolis effect are called the prevailing winds. In the tropics and   pressure  air moving from    Coriolis effect modifies this,   cyclones move swiftly over the ocean and can produce
            north-   westerlies  (Nov–Mar)  cold air flows   high to low pressure
          westerlies  easterlies  subtropics, the air movements toward the equator in Hadley cells are   toward area of low   deflected by Coriolis   producing circular winds.  rapid changes in wind strength and direction.  atmosphere influence one another.
        northeasterly   deflected to the west. These are known as trade winds. They   pressure  effect to form spiral  warm air cools    Most mollusks have a head, a soft body mass, and a muscular foot.
        trade winds  comprise the northeasterly trades in the Northern   at high altitude  cold air sinks    The foot is formed from the lower body surface and helps it to
                   Hemisphere, and southeasterly trades in the south.
                    At higher latitudes, the surface winds in Ferrel   Coastal Breezes  DAY AND NIGHT        move. Mollusks have what is called a hydrostatic skeleton—their
                                            Land heats up faster
                    cells deflect to the east, producing the westerlies.   Local winds, called onshore and offshore   than water during the day.
              Tropic of Cancer  In the Southern Hemisphere, these winds   breezes, are generated near coasts, especially in   air heats up   Warm air rises over the land   bodies are supported by internal fluid pressure rather than a hard
                                         and rises over  cool air
                    blow from west to east without meeting land.   sunny climes. Onshore breezes—sometimes   land  drawn in  and draws in cold air from   color-coded panel
                                            the sea. At night, the land
                    Those around latitudes of 40˚S are known as   called sea breezes—develop during the day.   skeleton. All mollusks have a mantle, a body layer that covers the
                                            reversing the airflow.
         Intertropical Convergence Zone  equator  the Roaring Forties. In polar regions, winds   These are caused by the land heating up more   cools more quickly,
                     deflect to the west as they move away from   quickly than the sea, as both absorb solar   ONSHORE BREEZE  shows position of   upper body and may or may not secrete a shell. The shell of
                     the poles. These are known as polar
       INTRODUCTION  southeasterly   southeasterly   KEY  prevailing   PATTERN OF WINDS  BREEZY COAST  small rise in temperature, whereas the same amount of heat energy is   cold air sinks cool air drawn   air heats up   INTRODUCTION  group being described   these can be held closed by powerful muscles while the tide is out,   REEF-DWELLING GOLIATH
                                  radiation. This occurs because the sea absorbs
                                                                                                             bivalves (clams and relatives) has two halves joined by a hinge;
                    northeasterlies and southeasterlies.
                                  large quantities of heat energy with only a
               Tropic of Capricorn
                                  likely to cause the land temperature to rise sharply (see p.31).
                                                                                                             or if danger threatens. Mollusks other than bivalves have a rasping
                       Year-round, the winds over
                                             and rises
                                  As the land warms up, it heats the air above it, causing the air to rise.
                     warm
                                            seaward
                                             over ocean
                       most oceans are trades or
                     prevailing
                                  Cooler air then blows in from the sea to take its place. In the evening,
                                                                                                             mouthpart, or radula, which is unique to mollusks. Cephalopods
                     cool
                       westerlies. An exception is the
                                  and at night, the opposite effect occurs. At nightfall, the land quickly
                       northern Indian Ocean—this
        trade winds
                   trade winds
                             On warm coasts, there is often a noticeable
                                  cools down, but the sea remains warm and continues to heat the air
                     local warm
                             drop in temperature from midday as a cool sea
                       has a monsoon climate, in
                                                                                                             (octopuses, squid, and cuttlefish) also have beaklike jaws as well
                       which a seasonal switch in
                             breeze blows in off the water. The breeze
                                  above it. As this warm air rises, it sucks the cooler air off the land, and
                     local cool
                       wind direction occurs.
                  southwesterly
              southeasterly
         westerlies
                  monsoon (Apr–Oct)
            southeasterlies
              trade winds
                westerlies
                                                                                                                            largest bivalve and may measure
                                                                                                             and snails) have a single shell. This is usually a spiral in snails,
                              WAVES AND TIDES
                             76 typically reverses in the evening and at night.  so generates an offshore breeze. This is sometimes called a “land breeze.”  OFFSHORE BREEZE  77  (indicated with    as tentacles, but most lack a shell, while most gastropods (slugs    The tropical giant clam is the
                                                                                                                            more than 3 ft (1 m) across and
                               Ocean Waves  Wave Generation  PLUNGING BREAKER    white outline) in the       but can be cone-shaped in other forms, such as limpets.   weigh over 440 lb (220 kg).
                                                    “Barrel” or “tube-forming” breakers like
                                        Wind energy is imparted to the sea surface through friction and
                                                    have large amounts of energy. The seabed
                                        pressure, causing waves. As the wind gains strength, the surface develops   this occur when the waves reaching shore
                               WAVES ARE DISTURBANCES in the ocean that transmit   gradually from flat and smooth through growing levels of roughness.   must be firm and quite steep.   gill
                               energy from one place to another. The most familiar   First, ripples form, then larger waves, called chop. The waves continue    classification hierarchy  spiral shell   mantle cavity  GASTROPOD ANATOMY
                                        to build, their maximum size depending on three factors: wind speed,
                               types of waves—the ones that cause boats to bob up   wind duration, and the area over which the wind is blowing, called    The body plan (far left)
                               and down on the open sea and dissipate as breakers    the fetch. When waves are as large as they can get under the current   of gastropods (slugs and
                               on beaches—are generated by wind on the ocean   conditions of wind speed and size of         snails) features a head,
                                        fetch, the sea surface is said to be “fully
                               surface. Other wave types include tsunamis, which    developed.”  The overall state of a sea surface   sensory   large foot, and usually
                               are often caused by underwater earthquakes (see p.49),   can be summarized by the significant wave   tentacle  a spiral shell (left). In
                                        height—defined as the average height of the
                               and internal waves, which travel underwater between    highest one-third of the waves. For example,   shelled forms, all the
                               water masses. Tides (see p.78) are also a type of wave.
                                        in a fully developed sea produced by winds
                                        of about 25 mph (40 kph), the significant                                           soft body parts can
                               Wave Properties  wave height is typically about 8 ft (2.5 m).  CAPILLARY WAVES (RIPPLES)  digestive   eye  be withdrawn into the
                                                                                                                            shell for protection, or
                                                                                                             system
                               A group of waves consists of several crests separated by troughs. The   disordered    These tiny waves are just a few
                                             millimetres high and have a
                                          sea surface
                               height of the waves is called the amplitude, the distance between   in fetch area            to conserve moisture
                               successive wave crests is known as the wavelength, and the time   ripples    outside the fetch,   wavelength of under 1½ in (4 cm).  SPIRAL    while uncovered by
                               between successive wave crests is the period. Waves are classified    turn to   chop  waves become   SNAIL SHELL
                                           sorted by speed
                               into types based on their periods. They range from ripples, which    wind   and wavelength    muscular foot   radula  the outgoing tide.
                               have periods of less than 0.5 seconds, up to tsunamis and tides,    direction   direction of    water motion occurs   wave shortens in length   wave finally
                                                                 breaks
                                                         offshore to depth of
                                                             and decreases in speed
                               whose periods are measured in minutes and hours (their    wave motion  half the wavelength   but increases in height
                               wavelengths range from hundreds to thousands of miles).                                  hinge
                               In between these extremes are chop and swell—the most
                               familiar types of surface wave. Ocean waves behave like                       BIVALVE ANATOMY  ligament   shell
                               light rays: they are reflected or refracted by obstacles they                 Bivalves are housed    mantle
                               encounter, such as islands. When different wave groups   CHOPPY SEA                           cavity
                               meet, they interfere—adding to, or canceling, each other.  In a choppy sea, the waves are      within a shell of two   digestive
                                             4–20 in (10–50 cm) high and have a
                                             wavelength of 10–40 ft (3–12 m).                                halves (right) from which   system
                                 wavelength    direction of                            GROUP INTRODUCTION ▶  the siphons and muscular
                               still-water   wave motion   BUILDING WAVES
                               level  crest    Within the wave-generation area,    direction  of wave        foot can be extended.
                                      the sea surface is usually quite
             TIDES AND WAVES ▶        confused—the result of groups    fetch (area    advance  wave reaches critical   water carried   The shell is opened and   muscular   siphon
                                      of waves of different size and
                                                                   up shore in
                                                              ratio of height to length
                                                                                                                      foot
                                          over which
                                                                                                             closed by the adductor
                                                                   swash zone
                                      wavelength interfering with each
                                          wind blows)
                                      other. Outside this area, the waves
                                      become sorted by speed to produce a   and begins to break   Pages such as the ones shown here describe   muscles, labeled in the   BIVALVE SHELL
                                      more regular pattern, called a swell.  FULLY DEVELOPED ROUGH SEA  water motion   Arrival on Shore  body plan (far right).    gill  adductor
                                                           caused by the
             The regular movements    PARTICLE MOVEMENT  trough    wave height    path of individual   Wind speeds over 40 mph (60 kph)    wave begins    As waves approach a shore, the motion they generate at depth begins    groups of organisms in general. All   muscle
                                             can generate very rough seas with
                                                           to interact
                                             waves more than 10 ft (3 m) high.
                                                             to interact with the sea floor. This slows the waves down and causes
                                   (amplitude)
                                    water particle
                                                           with the sea
                                                           bed and
                               As waves pass over the surface, the
                                                             the crests in a series of waves to bunch up—an effect called shoaling.
                               particles of water do not move forward   Wave Propagation  slow down  The period of the waves does not change, but they gain height as the
                               with the waves. Instead, they gyrate in   ROGUE WAVES  In the fetch, many different groups of waves of varying wavelength are   SHOALING AND BREAKING  energy each contains is compressed into a shorter horizontal distance,   jaws
                               little circles or loops. Underwater, the
            of the tides are described   particles move in ever-smaller loops. At   Interference between two or more large    generated and interfere. As they disperse away from the fetch, the waves   Shoaling occurs as waves enter   and eventually break.   radula  feeding arm
                                                       shallow water. The wave length and
                                   waves occasionally causes a giant or
                                                       speed both decrease, but the wave
                                                             There are two main types of breaker. Spilling breakers occur on flatter
                               a depth below about half the distance
                                        become more regularly sized and spaced. This is because the speed of
                                   “rogue” wave. This one, recorded in the
                                                       gains height. When the crest gets
                                                             shores: their crests break and cascade down the front as they draw near
                               between crests, they are quite still.
                                        a wave in open water is closely related to its wavelength. The different
                                   Atlantic Ocean in 1986, had an estimated   groups of waves move at different speeds and so are naturally sorted    too steep, it curls and breaks.  the shore, dissipating energy gradually. In a plunging breaker, which   introductions contain an account of the
                                   height of 56 ft (17 m). It broke over the ship                               eye
                                   pictured, bending its foremast back by 20˚.  by wavelength: the largest, fastest-moving waves at the fore, the smaller,   occurs on steeper shores,   digestive
                                                                  the crest curls and falls over
                                        slower-moving ones behind. This produces a regular wave pattern, or
             here, as well as the way   swell. Occasionally, groups of waves from separate storms interfere to   HUMAN IMPACT  the front of the advancing   defining physical characteristics, usually   system   arm
                                                                  wave, and the whole wave
                                                    RIDING THE WAVES
                                        produce unusually large “rogue” waves. As they propagate across the
                                                                  then collapses at once.
                                            open ocean, wind-generated
                                            waves maintain a constant speed,   When a swell reaches a suitably   Waves can also refract as   internal
                                            which is unaffected by depth    shaped beach, it can produce   they reach a coastline. This   shell  siphon  CEPHALOPOD ANATOMY
                                                    excellent surfing conditions.
            that disturbances spread        until they reach shallow water.   Small spilling breakers are ideal    concentrates wave energy   followed by further information on behavior,   OCEAN LIFE  gill  Cephalopods have large eyes, in front
                                                                  onto headlands (see p.93)
                                                    for novice surfers, while experts
                                            Only with waves of extremely
                                                    seek out large plunging breakers
                                            long wavelength—tsunamis—is
                                                                  and shapes some types of
                                                                                                                      of which there are a number of tentacles.
                                            the speed of propagation affected
                                                                  beach (see p.106).
                                                    that form a “tube” they can
                                                    ride along. For tube-riding, the
                                                                                                                      in rapid movement. Some forms have
                                                    smoothly either to the right or
            out across the surface in   INTRODUCTION  by water depth.  break of the wave must progress    WAVE REFRACTION  INTRODUCTION  habitats, and classification.  mantle cavity  The siphon functions in respiration and
                                            SWELL
                                                                                                                      a flattened internal shell.
                                                    left. Here, a surfer rides a right-
                                                                  When waves enter a bay enclosed
                                            A swell is a series of large, evenly
                                                    breaking wave in Hawaii—
                                                    it is breaking from left to right
                                            spaced waves, often observed hundreds
                                                                  by headlands, they are refracted
                                                                  (bent) as different parts of the
                                            of miles away from the storm that
                                            spawned them. Wavelengths range from
                                                    behind the surfer.
                                                                  wave-front encounter shallow
                the form of waves.          tens to hundreds of feet.   water and slow down.
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