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A.  Adaptation               B.  Differential threshold and luminance
                                       0.8
         10 6                          0.7
                  Rod
                  adaptation
         10 5                      vision  0.6
                   Cone          Day   0.5      Threshold
          4
                   adaptation
         10
        Relative luminance  10 3 2  Retinal  Dim-light  vision  Relative differential threshold ∆l/l  0.4  for day vision  Smallest  Adaptation to Light Intensity
                                       0.3
         10
         10
                                                         (0.01)
                                       0.1
          1  adaptation                0.2           differential threshold
            Absolute threshold
                 10     20     30           10 4  10 6  10 8  10 10
                   Adaptation time (min)         Relative luminance
                                                         (After G. Schubert)
         C.  Mechanisms for adaptation                                 Plate 12.22
          Dim light      Bright light    Dim light       Bright light
                 1   Pupil reflex             2  Sensory proteins







          Wide pupil     Narrow pupil      Ample           Sparse
                                             Rhodopsin and transducin
               3  Spatial Summation         4  Temporal Summation

                                         Dim light     Bright light
                                        Potential


                                              Time           Time
         Large retinal area  Small retinal area
         for each neuron  for each neuron  Longer stimulus  Brief stimulus
                                          required for AP  sufficient for AP
         D.  Successive contrast (“local adaptation”) (see text)





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