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Physical Fitness and Training   types of physical training strategies, and most
                                       training programs use a combination of them.
       The  physical  exercise  capacity  can  be  Motor learning, which increases the rate and
       measured using simple yet standardized tech-  accuracy of motor skills (e.g., typewriting).
       niques of ergometry. This may be desirable in  These activities primarily involve the CNS.
       athletes, for example, to assess the results of  Endurance training, which improves sub-
       training, or in patients undergoing rehabilita-  maximal long-term performance (e.g., run-
       tion therapy. Ergometry assesses the effects of  ning a marathon). The main objectives of en-
       exercise on physiological parameters such as  durance training are to increase the oxidative
                   .
    Nerve and Muscle, Physical Work  (performance) is expressed in watts (W) or  to increase V O 2 max (! B, C). The resulting in-
                                       capacity of slow-twitch motor units (! p. 58!,
       O 2 consumption (V O 2 ), respiration rate, heart
                                       e.g., by increasing the mitochondrial density,
       rate (! p. 74), and the plasma lactate concen-
       tration (! A). The measured physical power
                                       increase the cardiac output and, consequently,
                                              .
                                       crease in heart weight allows higher stroke
       W/kg body weight (BW).
                                       volumes (! C) as well as higher tidal volumes,
       In bicycle ergometry, a brake is used to adjust the watt
                                       resulting in very low resting heart rates and
       level. In “uphill” ergometry on a treadmill set at an
                                       respiratory rates. Trained athletes can there-
       angle α, exercise performance in watts is calculated
                                       fore achieve larger increases in cardiac output
       as a factor of body mass (kg)!gravitational accelera-
                                       and ventilation than untrained subjects (! C).
              – 2
       tion g (m · s )!distance traveled (m)!sin α!1/
                                          .
       time required (s ). In the Margaria step test, the test
                – 1
                                       by the cardiovascular capacity, not the respira-
       possible after a certain starting distance. Perform-
                                       tory capacity. In individuals who practice en-
       ance is then measured as body mass (kg)!g
    2  subject is required to run up a staircase as fast as  The V O 2 max of a healthy individual is limited
                                       durance training, the exercise-related rise in
          – 2
                       – 1
       (m · s )!height/time (m · s ).
                                       the lactate concentraton is also lower and oc-
       Short-term  performance  tests  (10–30 s)  curs later than in untrained subjects (! A).
       measure performance achieved through the  Strength training improves the maximum
       rapidly available energy reserves (creatine  short-term performance level (e.g., in weight
       phosphate, glycogen). Medium-term perform-  lifting). The main objectives are to increase the
       ance tests measure performance fueled by an-  muscle mass by increasing the size of the
       aerobic glycolysis (! p. 72). The maximum O 2  muscle fibers (hypertrophy) and to increase
                 .
       consumption (V O 2 max) is used to measure  the glycolytic capacity of type motor units
       longer term aerobic exercise performance  (! p. 58).
       achieved through oxidation of glucose and free  Excessive physical exercise causes muscle
       fatty acids (! p. 74).          soreness and stiffness. The underlying cause is
         In strenuous exercise (roughly 2/3 the max-  not lactic acid accumulation, but sarcomere
       imum physical capacity or more), the aerobic  microtrauma, which leads to muscle swelling
       mechanisms do not produce enough energy, so  and pain. The muscle ache, is a sign of micro-
       anaerobic metabolism must continue as a par-  inflammation (! D).
       allel energy source. This results in lactacidosis  Muscle fatigue may be peripheral or central.
       and a sharp increase in the plasma lactate con-  Peripheral fatigue ist caused by the exhaustion
       centration (! A). Lactate concentrations of  of energy reserves and the accumulation of
       up to 2 mmol/L (aerobic threshold) can be  metabolic products in the active muscle. This
       tolerated for prolonged periods of exercise.  is particularly quick to occur during postural
       Lactate concentrations above 4 mmol/L (an-  work (! p. 66). Central fatigue is characterized
       aerobic threshold) indicate that the perform-  by work-related pain in the involved muscles
       ance limit will soon be reached. Exercise must  and joints that prevents the continuation of
       eventually be interrupted, not because of the  physical exercise or decreased the individual’s
       increasing lactate concentration, but because  motivation to continue the exercise.
       of the increasing level of acidosis (! p. 74).
         Physical training raises and maintains the
   76  physical exercise capacity. There are three
       Despopoulos, Color Atlas of Physiology © 2003 Thieme
       All rights reserved. Usage subject to terms and conditions of license.
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