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PLANTS

       Stems                                                             MICROGRAPH OF LONGITUDINAL
                                                                         SECTION THROUGH APEX OF STEM
                                                                         Coleus sp.

       THE STEM IS THE MAIN SUPPORTIVE PART OF A PLANT that grows
       above ground. Stems bear leaves (organs of photosynthesis), which    Apical
                                                                        meristem            Procambial
       grow at nodes; buds (shoots covered by protective scales), which grow    (region of    strand (cells
       at the stem tip (apical or terminal buds) and in the angle between a   actively      that produce
       leaf and the stem (axillary or lateral buds); and flowers (reproductive   dividing   vascular tissue)
                                                                        cells)
       structures). The stem forms part of the plant’s transport system: xylem               Leaf  primordium
       tissue in the stem transports water and minerals from the roots to the                (developing leaf)
       aerial parts of the plant, and phloem tissue transports nutrients
                                                                    Developing               Cortex
       manufactured in the leaves to other parts of the plant. Stem tissues   bud            (layer between
       are also used for storing water and food. Herbaceous (non-woody)                      epidermis and
                                                                                             vascular tissue)
       stems have an outer protective epidermis covering a cortex that
       consists mainly of parenchyma (packing tissue) but also has some                     Vascular
       collenchyma (supporting tissue). The vascular tissue of such stems                   tissue
       is arranged in bundles, each of which consists of xylem, phloem,                     Epidermis
       and sclerenchyma (strengthening tissue). Woody stems have an outer   Pith            (outer layer
       protective layer of tough bark, which is perforated with lenticels (pores)           of  cells)
       to allow gas exchange. Inside the bark is a ring of secondary phloem,                         Young
       which surrounds an inner core of secondary xylem.                                             leaves
                                                                                                     emerging
       YOUNG WOODY STEM                                                  EMERGENT BUDS
       Lime                                                              London plane
       (Tilia sp.)                                                       (Platanus x acerifolia)

                            Secondary     Pith               Phellem
                            phloem                           (protective          Terminal bud
         Cortex                                              cork layer)
         (layer between phellem
         and vascular tissue)

        Xylem vessel
        (through which                                                      Lateral bud
        water and
        minerals are
        transported)                                             Vascular cambium
                                                                 (actively dividing cells            Node
          Xylem fiber                                            that produce xylem
          (supporting                                            and phloem)                     Internode
          tissue)
                                                                            Inner
          Ray                                                   Fall        bud scale
          (parenchyma                                           wood
          cells)                                                        Secondary
                                                                Spring   xylem
                                                                wood
       Phloem sieve tube                                                     Outer
       (through which                                                        bud scale
       nutrients are
       transported)                                                               Node            Leaf  scar
                                                                 Companion cell
                                                                 (cell associated
          Phloem fiber                                           with phloem
          (supporting                                            sieve tube)
          tissue)                                                                Lenticel
                                                                                 (pore)           Woody
               Lenticel                                                                           stem
               (pore)


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